The Dr. Jonathan Waters Waters is a biologist at the University of Otago, New Zealand. He specialises in the analysis of genetic variation in spatial and evolutionary contexts. His marine biological research on marine mammals, penguins, kelp and invertebrates emphasises the roles of extinction and long-distance dispersal in shaping biological assemblages of the Southern Hemisphere. His ancient-DNA research has highlighted dramatic prehistoric shifts in several of New Zealand's iconic coastal vertebrate species. His freshwater fish and insect research highlights the tight relationships between tectonic processes and biological evolution..